فهرست:
چکیده
فصل اول
مقدمه ................................................................................................................................................................ 1
1- بررسی منابع .................................................................................................................................................. 3
1-1- تاریخچه کنترل علفهای هرز .................................................................................................................. 3
1-2- پراکنش و اهمیت علف هرز ..................................................................................................................... 4
1-3- تعاریف علف هرز .................................................................................................................................... 6
1-4- خصوصیات علف هرز .............................................................................................................................. 7
1-5- خسارتهای علف هرز ............................................................................................................................. 8
1-5-1- خسارتهای کمی ................................................................................................................................ 8
1-5-2- خسارتهای کیفی ............................................................................................................................... 9
1-6- کنترل علفهای هرز ................................................................................................................................... 9
1-7- گیاهشناسی یولاف وحشی ...................................................................................................................... 10
1-8- گیاهشناسی بارهنگ ................................................................................................................................ 10
1-9- اکولوژی یولاف .................................................................................................................................... 11
1-10- مراحل رشد یولاف .............................................................................................................................. 12
1-11- مرحله جوانه زنی .................................................................................................................................. 13
1-12 ساختمان دانه .......................................................................................................................................... 13
1-13- بیوشیمی و فیزیولوژی جوانه بذر ........................................................................................................... 15
1-14- کنترل علفهای هرز ............................................................................................................................... 15
1-15- دلایل غالبیت بذور علفهای هرز ........................................................................................................... 16
1-15-1- پوسته بذر ........................................................................................................................................ 16
1-16- خواب بعد از رسیدگی ......................................................................................................................... 17
1-17- تعریف خواب ...................................................................................................................................... 18
1-18- طبقهبندی خواب بذر ............................................................................................................................ 18
1-19- اهمیت خواب بذر ................................................................................................................................ 20
1-20- بررسی انواع خواب .............................................................................................................................. 20
1-20-1- خواب فیزیولوژی ............................................................................................................................ 20
1-20-2 خواب مورفولوژیکی ......................................................................................................................... 21
1-20-3- خواب فیزیکی ................................................................................................................................. 21
1-20-4- خواب شیمیایی ................................................................................................................................ 21
1-20-5- خواب مکانیکی ............................................................................................................................... 22
1-21- اهمیت خواب در کشاورزی کاربردی .................................................................................................. 22
1-21-1 خواب در گیاهان زراعی و ارتباط با قوه نامیه ...................................................................................... 22
1-21-2- خواب یا مرگ بذر؟ ........................................................................................................................ 22
1-22- فاکتورهای که باعث ایجاد خواب در بذر میشوند ................................................................................ 23
1-23- پوشش ایجاد کننده خواب بذر ............................................................................................................. 23
1-24- نقش هورمونها و تنظیم کنندههای رشد در خواب بذر ........................................................................... 25
1-24-1- بازدارندههای جوانهزنی .................................................................................................................... 25
1-24-2 تولیدهکنندههای جوانهزنی .................................................................................................................. 26
1-25- روشهای برطرف کردن خواب .............................................................................................................. 27
1-26- شکستن خواب یا به کارگیری مواد شیمیایی ......................................................................................... 28
1-27- مکانیسمهای موثر در برطرف شدن خواب جنین به وسیله خواب کنترل کننده ....................................... 29
1-28- نیازهای خاص در آزمون جوانهزنی ....................................................................................................... 30
1-29- ارزیابی آزمون جوانهزنی ...................................................................................................................... 31
1-30- آزمونهای بنیه که براساس تاثیرات فیزیولوژیکی پیری است ................................................................. 32
1-31- آزمونهای بنیه که براساس تاثیرات بیوشیمیایی در طی پیری است ......................................................... 33
فصل دوم ......................................................................................................................................................... 39
2- مواد و روشها .............................................................................................................................................. 40
2-1- مراحل اجرای آزمایش ........................................................................................................................... 40
2-2- طرح آزمایشی ........................................................................................................................................ 40
2-3- تیمارهای آزمایشی ................................................................................................................................. 41
2-4- صفات مورد مطالعه ................................................................................................................................ 42
2-5- تجزیه آماری .......................................................................................................................................... 43
فصل سوم ........................................................................................................................................................ 44
3- نتایج و بحث ............................................................................................................................................... 45
3-1- تجزیه واریانسی صفات مورد مطالعه گیاه یولاف در تیمار با نیترات پتاسیم ............................................... 45
3-2- مقایسات میانگین صفات مورد مطالعه در تیمار نیترات پتاسیم ................................................................... 47
3-3- مقایسه میانگین اثر تیمار اسید جیبرلیک در گیاه یولاف ............................................................................53
3-4- مقایسات میانگین صفات مورد مطالعه در تیمار اسید جیبرلیک ................................................................. 55
3-5- نتیجهگیری کلی ..................................................................................................................................... 62
3-6- پیشنهادات .............................................................................................................................................. 63
فهرست منابع فارسی ......................................................................................................................................... 64
فهرست منابع غیر فارسی ................................................................................................................................... 67
چکیده به انگلیسی ............................................................................................................................................ 72
منبع:
Agrawal pk .1992. Germination tests under controlled conditions and its Evalution. In pk Agrawal, M Dalani, eds, Techniques in seed science and technology. International Book Company, N. Y., pp 61-84.
Albrecht, K. A., E. A. oelke, and M. L. Brenner. 1979. Crop sci. 19:671-76.
Amen, R. 1963. Am. Sci. 51:408-24.
Anonymous. 1993. ISTA- International Rules for seed testing, supplement 21.
Arias. L. 1976. Studies in seed dormancy. IX. The vole of gibberellins biosynthesis and the release of gibberellins in seeds of corylus arellana L. plant a. 131:135-139.
Arios, I. Williams, P. M. and Bradbeer, J. W. (1926) studies in seed dormancy. Ix. The vole of gibberellins biosynthesis and the release of bound gibberellins in the post- chilling accumulation of gibberellins in seeds of corylus avellana L. planta (Berlin) 131, 135-139.
Arteca, R. N. 1996. Plant Gorowth substances principles and Application chapter 3: chemistry, Biological effects and Mechanism of Action, p: 66.
Baskin, C. C. and Baskin, Y. M. 2004. A Classification system for seed dormancy. Seed science Research 14:1-16.
Basra, A. S. 1994. Seed quality basic mechanism and agricultural implications. Food products press. P: 386.
Bayer, A. G. 1986. Important crops of the world and their weeds.
Benech, R. A. Sanchez, f. Forcella. 2000. Environmental control of dormancy in weed seed banks in soil. Field crops Research 67:105-122.
Bewlcy, Y. D. and Black, M. 1994. Seeds: physiology of development and germination plenum press, New York.
Black, Y. N. 1956. Aust. Y. Agric. Res. 7:98-109.
Borriss, H. 1949. Jahrb. Wiss. Bot. 89:254-339.
Brad ford, K. Y., Y. Y. Steiner, and S. E. Trowatha. 1990. Seed priming influence on germination and emergence of pepper seed lots. Crop seience 30:708-721.
Bradford. K. Y. 2002. Applications of hydrothermal time to quantifying and modeling seed germination and dormancy. Weed science 50:248-260.
Brothwrck, H. A., S. B. Hendricks, E. H. Toole, and N. K. Toole. 1954. Bot. Gaz. 115:205-25.
Burris, Y. S., O. T. Edge, and A. H. Wahab. 1973. Crop sci. 13:207-10.
Ching, T. M., and W. H. Foote. 1961. Agron. Y. 53:183-86.
Clark, B. E., and N. H. peck. 1968. N.Y. Agric. Exp. Stn Bull. 819.
conins, A. L. P. and Devililliers, O. T. 1986. Breaking of Arena Fatua L. seed by treatment with ammonia. Weed Res. 26:191-197.
Copeland, L. O. 1967. Principles of seed science and Technology. Minneapolis: Burgess.
Esno H .. Solna H .. and Sweden M. 1996. Proceeding of the International Seed T esting AssociationWageningen؛ Thr Netheriands.P.92.
Faster, S. and J. A. Duke. 1990. Medicinal plants. The peterson field Guide series, Houghton Mifflin company, 366 p, New York.
Finch, W. E. 2006. Seed dormancy and the control of germination. New phytologist. 171:501-523.
Forcella, F. Bencch Arnold, R. L., Sanchez, R. and chersa, C. M. 2000. Modeling seeding emergence field, crops Res. 67:123-139.
Fracoise, F. A. 1998. Effect of cinnamic acid on poly phenol production in Iplantago Lanceolata. Phytochem. 49:697-702.
Hampton YR .1979. An Introduction to seed Technology. Halstead press, New York.
Hay, y.R. 1967. In physiologic, okologic, und Biochemic der kemiung, I, ed. H. Borriss. Grefswald, Grnst- Moritz-Arndt University at.
Hilton, Y. R. 1984. The in fluencies of light and potassium nitrate on the dormancy and germination of Arena fatua L. seed and its ecological significance. New phytol. 96:31-34.
Hit horst, H. W. 1995. A critical update on seed dormancy. I. primary dormancy. Seed sci. Res., 5:61-73.
Hsiao, A. I. 1985. Wild oats (Avena Fatua L.) Seed dormancy as in fluenced by sodium hypochlorite, moist storage and gibberellins A3. Weed Res. 25:281-288.
Hsiao, A. I., and Qulck, W. A. 1985. Wild oats (Avena Fatual.) seed dormancy as influenced by sodium hypocworite, moist storage and gibberellins A3. Weed Res. 25:281-288.
Ipek, A. 2005. Role of GA and KNO in improving the frequency of seed Germination in plantago lanceolatall. 37:883-887.
Karssen, C. M. 1982. Seasonal patterns of dormancy in weed seeds. In: “the physiology and Biochemistry of seed. Dormancy and Germination” (A. A. Khan, E. d.). pp. 243-270, Elsevier, Amsterdam.
Karssen, C. M. and R. weges. 1989. Physiological mechanisms involved in seed priming In: Recent Advances in the development and Germination. Blackwell publishing (volume 27).
Kavssen, C. M. 1983. Induction of dormancy during seed development by endogenous abscisic acid: studies on abscisic acid deficient genotypes of Arabidopsis thaliana heynh. 157:158-65.
Khan, A. A. 1977. In: the physiology and Biochemistry of seed Dormancy and germination, pp. 29-50. North Holland publishing co., Amsterdam.
Kondo, T. 1993. Promotion of hard- seed germination in lotus corniculatus var Japonica for use in amenity grasslands. Seed science and Technology, 21:611-619.
Kucera, B. 2005. Plant hormone in teraetions during seed dormancy release and germination. Seed science Research 15:281-307.
Leon, R. G. and knap, A. D. 2004. Effect of temperature on the germination of common water hemp (Amaran thus tuberc ulatus). Gaint Foxtail (setaria Faberi), and velbet leaf (abutilon the ophrasti). Weed sci 52:67-73.
Leopold, A. C. 1975. Plant Growth and Development. NewLeubner, G. 2003. Functions and regulation of glucanase during seed germination, dormancy release and after- ripening. Seed science Research 13:17-34.
Lyshede, O. B. 1992. Studies on mature seeds of cuscuta pedicellat and C. campestris by
electron microscopy. Ann. Bot. 69:365-371.
Maguire J.D..and Overiand A. 1959 . Laboratory germination of seed of weedy and native plants Wash.
Meyers, O., G. Gaffney, and D. Hall.1979. Abstracts Ill. State Acad. Sci.
Miller, C. O. 1956. Effects of cytokinings on germination of lettuce seeds. Plant physiol.,
Myers, S. P. 1997. Developmental differences between germination after- ripened and dormant excised Arena Fatua L. embryos. Ann. Bot. 79:10-23.
Ni, B. R. 1993. Germination and dormancy of abscisic acid and gibberellins- deficient mutant tomato seeds. Sensitivity of germination to abscisic acid, gibberellins, and water potential. Plant physiology 101:607-617.
Nicois MA, Heydecker W .1968. “Tow approaches to the study of germination data.” Proc. Int. seed test. Asso. 33:531-540.
Nikolaeva, M. G. 1977. Factors controlling the seed dormancy pattern. In “The physiology and Bio chemistry of seed dormancy and Germination” pp. 51-74. North- Holland.
Nikolaeva, M. G. 1977.Factors controlling the seed dormancy pattern. In “the physiology and Biochemistry of seed Dormancy and Grmination” (A. A. Khan, ed.), pp. 51-74. North- Holland, Amsterdam/ New York.
Odum, S. 1965. Germination of ancient seads. Dan- Bot. Ark. 24:1-70.
Omami, EN. Medd. RW. 1992. Germination and after-ripening responses in Amaranthus rctor flexus seed. Proceeding of the first.
Ooi, M. K. 2006. Dormancy and the Five- centric focus: germination of three leucopogon species from south- eastern Australia. Annals of Botany 98:421-43.
Ozaga, J. A. 1992. Seed effects on gibberellins metabolism in pea pericarp. Plant physiol. 100:88-94.
Pollock, B. M. and Toole V. K. 1961. After ripening, rest period and dormancy.
Reisman, O. J. Kigel. And B. Rabin. 1991. Dormancy patterns in buried seeds of Datura Sorox and Datura stramonium. Can. J. Bot. 69:173-179.
Roberts, E. H. 1972. Storage environment and the control viability. In viability of seeds, ed. E. H. Roberts, Chapman and Hall. London, 14-58.
sanderson, M. A. and G. F. Elwinger. 2000. Chicory and English plantain seedling emergence at different planting depths. Agronomy Y., 92:1206-10.
Saruhan, N. 2002. Alleviation of seed dormancy in plant ago major. Israel Y. plant sci. 50:177-9.
Simpson, G.M. 1978. n Dormancy and Development Arrest, ed. M. E. cutter. New.
Toole, E. H., and S. Hendricks. 1956. Annu. Rev. Plant physiol. 7:229-324.
Totterdell and E. H. Roberts. 1979. Effect of low temperatures on the loss of innate dormancy and development of induced dormancy in seeds of Rumex obtusifolius L. and Rumex Crispus L. plant, cell and Environment 2:131-137.
Vivrette, N. 2001. Seed dormancy (chapt. 9) In: seed Technologist Training Manual; soc. Of comm. Seed Technologists; M. MC Donald, T. Gutormson, B. Turnip seed.
Welbaum, G. E. 1990. Water relationof seed development and germination in muskmelon (cucumis melo L.). Plant physiol. 92:1029-1037.
Yacobsen, Y. V. 1976. The proteins velased by isolated barley aleurone alyers before and after gibberllic acid treatment. Plant a. 115:193-206.
Baskin. C. 1986. Eco physiology of secondary dormancy in seeds of Ambrosia artemisiifoli . Ecology 61:472-480.
Hillman, W. S. 1962. The physiology of flowering. New York: Holt, Rinehart, and Winston.
Watkins. J. T. 1983. Mechanical resistance of the seed coot and endosperm during germination of capsicum annum at low temperature